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Montana Field Guides

Low Hairy Goldenaster - Heterotheca villosa var. depressa
Other Names:  Chrysopsis depressa, Heterotheca depressa

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Native Species

Global Rank: G5T3
State Rank: SU
(see State Rank Reason below)
C-value:


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State Rank Reason (see State Rank above)
Heterotheca villosa var. depressa is described by Semple (1996) in his publication, A Revision of Heterotheca sect. Phyllotheca (Nutt.) Harms (Compositate; Astereae): The Prairie and Montane Goldenasters of North America and treatment in the Flora of North America (FNA; Semple in FNA 2006). Variety depressa occurs in Yellowstone National Park and adjacent locations (Semple 1996). Semple found that duplicate specimens collected near Gardiner, Montana by Hitchcock and Muhlick (#13577) in 1945 and deposited at six herbaria contained plants representing varieties depressa and minor along with intermediary forms (1996). Semple's 2006 treatment published in FNA only credits Wyoming as having variety depressa, and states that varieties depressa and minor can hybridize. Since then Booth's 1941 collection (ID 139919) from near Livingston, Montana has been annotated as Heterotheca depressa by Nesom in 2017. A conservation status rank of unrankable (SU) is assigned in Montana because this variety appears to be present in Montana, yet intermediate forms can exist, specimen locations are poorly mapped, and current information is lacking.
 
General Description
PLANTS: Perennial plants that grow from a taproot. Stems decumbent to erect, woody at the base, and 10–50 cm tall. Hairs are strigose (stiff, straight, sharp, and appressed) to hirsute (long and stiff), often stipitate-glandular (stalked glands). Source: Lesica et al. 2012

LEAVES: Alternately arranged on the stem (cauline). Those on the lower stem petiolate and upwards becoming sessile. Leaf blades are oblanceolate to obovate in shape with smooth (entire) margins, and 1–5 cm long. Source: Lesica et al. 2012

INFLORESCENCE: Yellow flowers in heads arranged in leafy-bracted corymbiform arrays. Flower stems (peduncles) are 0–3 cm long. Involucre is narrowly campanulate, 5–10 mm high. Involucral bracts (phyllaries) are imbricate forming 4 to 5 rows; bracts are green, narrowly lanceolate, white-margined, and glabrate to strigose, though sometimes glandular. The receptacle is nearly flat. Heads composed of yellow ray and disk florets (radiate). Fruit is a Cypsela, a specific type of achene. Source: Lesica et al. 2012

VARIETY depressa: Stems are 5-22 cm tall and sparsely long-hirsute. Upper Leaves are densely strigose (84-140 hairs per square millimeter) and sparsely to moderately stipitate-glandular. Typical Habitats are hot springs and gravelly stream banks. Source: (Semple in FNA 2006)

Heterotheca is derived from the Greek words heteros meaning 'different' and thece meaning 'container' which refers to ray and disk cypsela being morphological different in some species (dimorphic cypsela) (Semple in Flora of North America [FNA] 2006).

Diagnostic Characteristics
Heterotheca villosa has 5 varieties in Montana (Semple in FNA 2006), of which 3 varieties are recognized in the Manual of Montana Vascular Plants (Lesica et al. 2012):

Variety depressa
Flora of North America
*Overall: Small stature, small flower heads, and linear-oblanceolate leaves with many hairs and usually some glands.
*Stems: 5-22 cm tall and sparsely long-hirsute.
*Upper Leaves are densely strigose (84-140 hairs per square millimeter) and sparsely to moderately stipitate-glandular.
*Distribution: Yellowstone National Park and adjacent areas.

Variety ballardii
Flora of North America
*Overall: Plants have leaves with round bases, short hairs, and not glandular and flower heads that are large, many-rayed and often subtended by one or more large, narrow, leaf-like peduncle bracts.
*Stems: 22-50 cm tall and usually abundantly long-hirsute.
*Flower Heads: Broadly campanulate involucre. Showy ray florets, (12)17-20 in number, (8.5)9-14.5 mm in length.
*Distribution: Northeastern Great Plains.

Variety foliosa
Flora of North America
*Stems: 5-22 cm tall and usually abundantly long-hirsute.
*Flower Heads: Narrowly campanulate involucre. Less showy ray florets, (7)13-22 in number, 8-11.4(12.4) mm in length.
*Distribution: Mountains and western prairies.
Manual of Montana Vascular Plants
*Stems and upper leaves densely hirsute and lacking glands.
*Flower heads on very short peduncles and immediately subtended by leaves.
*Leaves are oblong to obovate in shape.
*Found mostly along streams on the plains in Montana.

Variety minor
Flora of North America
*Overall: Upper leaves are narrowly to broadly oblanceolate (sometimes oblong or ovate in the mountains), moderately glandular (4-20 glands per square millimeter), and hispido-strigose (10-50 hairs per square millimeter). Upper leaf bases are cuneate or attenuate, but rarely rounded.
*Stems: decumbent to ascending-erect, (9)16-33(48) cm tall. Branches usually not very long.
*Upper Leaves: Margins flat to undulate with obtuse to acute leaf tips, and sparse to moderate hairs.
*Flower Heads: Arrays are compact. Involucral bracts are moderately strigose (hairs to 1 mm long) and usually stipitate-glandular.
*Distribution: Throughout most of the range described for the species.
Manual of Montana Vascular Plants
*Stems sparsely to densely stipitate-glandular, but not densely hirsute.
*Found in plains, valleys, to rarely the subalpine in Montana.
*Montana's most common form.

Variety villosa
Flora of North America
*Stems: 16-39 cm tall and moderately to densely strigose and sparsely to abundantly long-hirsute.
*Upper Leaves are moderately strigose (20-60 hairs per square millimeter) and sparsely to moderately stipitate-glandular.
*Flower Heads: Usually lack bracts on the peduncles.
*Distribution: Great Plains and scattered in hills and mountains.
Manual of Montana Vascular Plants
*Stems and upper leaves densely hirsute and lacking glands.
*Flower heads on longer peduncles and subtended by small bracts.
*Leaves are more oblanceolate in shape.
*Found mostly on the plains in Montana.

Range Comments
Wyoming (Semple in FNA 2006) and evidently Montana (see State Rank Reason).

Observations in Montana Natural Heritage Program Database
Number of Observations: 2

(Click on the following maps and charts to see full sized version) Map Help and Descriptions
Relative Density

Recency

 

(Observations spanning multiple months or years are excluded from time charts)



Habitat
Hot springs, broken travertine on old terraces, dry geyserite, gravelly stream banks, dry rock hillsides, open slopes, and dry sandy soils in montane conifer forests; 1,800-2,400 meters in elevation (Semple in FNA 2006). Where in vicinity of each other Variety depressa is typically present in vicinity of hot springs, geyser basins, and nearby river flood banks while variety minor occupies drier or non-geyserite habitats.

Reproductive Characteristics
Varieties depressa and minor can hybridize (Semple in FNA 2006).

Management
Variety depressa is of conservation concern in Wyoming (Semple in FNA 2006).


References
  • Literature Cited AboveLegend:   View Online Publication
    • Flora of North America Editorial Committee. 2006. Flora of North America North of Mexico. Vol. 20. Magnoliophyta: Asteridae, part 7: Asteraceae, part 2. Oxford Univ. Press, New York. xxii + 666 pp.
    • Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p.
    • Semple, J.C. 1996. A Revision of Heterotheca Sect. Phyllotheca (Nutt.) Harms (Compositae: Astereae): the Prairie and Montane Goldenasters of North America. University of Waterloo Biology Series 37: 1-164.
  • Additional ReferencesLegend:   View Online Publication
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    • Durham, R. A., D. L. Mummey, L. Shreading, and P.W. Ramsey. 2017. Phenological patterns differ between exotic and native plants: Field observations from the Sapphire Mountains, Montana. Natural Areas Journal, 37(3), 361–381.
    • King, L.A. 1980. Effects of topsoiling and other reclamation practices on nonseeded species establishment on surface mined land at Colstrip, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 129 p.
    • Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p.
    • Meier, G.A. 1997. The colonization of Montana roadsides by native and exotic plants. M.Sc. Thesis. Bozeman, MT: Montana State University. 45 p.
    • Rennick, R.B. 1981. Effects of prescribed burning on mixed prairie vegetation in southeastern Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 144 p.
    • Sater, S. 2022. The insects of Sevenmile Creek, a pictorial guide to their diversity and ecology. Undergraduate Thesis. Helena, MT: Carroll College. 242 p.
    • Simanonok, M.P. and L.A. Burkle. 2019. Nesting success of wood-cavity-nesting bees declines with increasing time since wildfire. Ecology and Evolution 9:12436-12445.
  • Web Search Engines for Articles on "Low Hairy Goldenaster"
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Low Hairy Goldenaster — Heterotheca villosa var. depressa.  Montana Field Guide.  .  Retrieved on , from